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Here's an incredibly lucky, rare look at courtship in wild cougars. F108 is the female without a collar. Here she is caterwauling, moaning, growling and rubbing her mate, M85, an adult male with a collar. Females require physical contact to force an egg to drop in order to facilitate fertilization and pregnancy. Thus all of the rubbing! This is representative of every mating we've seen--the female typically initiates and convinces the male to participate, more than the other way around.